HERA was a particle accelerator at DESY in Hamburg. It was operated from 1992 to 30 June 2007. At HERA, electrons or positrons were brought to collision with protons at a center-of-mass energy of 320 GeV. HERA was used mainly to study the structure of protons and the properties of quarks, laying the foundation for much of the science done at the Large Hadron Collider (LHC) at the CERN particle physics laboratory today. HERA is the only lepton–proton collider in the world to date and was on the energy frontier in certain regions of the kinematic range.